This Pseudocordylus+ site includes also the genus Hemicordylus.

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1838 Pseudocordylus Smith
1838 Hemicordylus Smith
2011 Pseudocordylus – Stanley et al.
2011 Hemicordylus – Stanley et al. (type species: Cordylus (Hemicordylus) capensis Smith 1838)


Smith, A. (1838): Contributions to the South African Zoology. — Annals and Magazine of Natural History (London), (2) 2: 31–34. [type species: Pseudocordylus montanus Smith, 1838 (= Pseudocordylus microlepidotus microlepidotus (Cuvier, 1829))]

Distribution: Rep. South Africa (excluding most of the Kalahari and south-east coastal zone). [Lang 1991: map]

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The new Cordylidae systematic – from Stanley, Bauer, Jackman, Branch & Mouton (2011) –
was included in parts; for more details
see here.
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Pseudocordylus capensis (Smith, 1838) (Hemicordylus capensis (Smith, 1838 [Stanley et al. 2011])
1838 Cordylus (Hemicordylus) capensis Smith
1839 Zonurus capensis – Duméril & Bibron
1921 Zonurus robertsi van Dam
1943 Cordylus capensis capensis – Fitzsimons
1943 Cordylus capensis robertsi – Fitzsimons
1944 Pseudocordylus robertsi – Loveridge
1992 Pseudocordylus capensis – Herselman, Mouton & van Wyk
2001 Cordylus capensis – Frost, Janies, Mouton & Titus
2002 Cordylus capensis – Anonymus/CITES

Smith, A. (1838): Contributions to the South African Zoology. — Annals and Magazine of Natural History (London), (2) 2: 31–34. [as Cordylus (Hemicordylus) capensis; type locality: “Cape of Good Hope”; type in NHM]

Pseudocordylus capensis  Fitzsimons, 1943  Smith, 1838-1849
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Anonymus/CITES (2002): distr  • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Boulenger 1885: morph •  Branch 1981: distr, behav  • Branch 1998: morph, breeding, biology, habitat, distr, map •  Branch & Bauer 1995: behav •  Cooper, van Wyk, Mouton, Al-Johany et al. 2000: antipredatory behavior  • Cooper, Whiting & van Wyk 1997: foraging modes  • Curtin, Mouton & Chinsamy 2005: bone growth patterns • van Dam 1921: description of Zonurus robertsi (type locality: “Klaver, van Rhynsdorp Dist., Cape Province”, type in TM)  • Eifler, Eifler & Eifler 2007: habitat use, behaviour • Fitzsimons 1935[a]: morph, colour •  Fitzsimons 1937: type in NHM •  Fitzsimons 1943: sys, morph, colour, size, distr, behav • Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces  • Herselman, Mouton & van Wyk 1992[a]: sys, distr, morph •  Hewitt 1937[a]: morph •  Hughes 1988: age: >3 years •  Loveridge 1944: revision (=all data)  • Mouton 1986[b]: distr •  Mouton & van Wyk 1989[a]: sys •  Power 1930: morph • Rensburg & Mouton 2009: foraging mode, space use, territoriality •  Rogner 1994: morph, behav, ter •  Smith 1838-1849 [= 1843]: morph, plate 27 (1843) fig. 2: Cordylus (Hemicordylus) capensis = Pseudocordylus capensis •  Visser 1984[d]: morph, behav, distr •  Wermuth 1968: sys, distr  • van Wyk & Mouton 1998: reproduction and sexual dimorphism  • van Wyk & Mouton 1992: epidermal glandular structure •


Pseudocordylus langi Loveridge, 1944
1944 Pseudocordylus langi Loveridge
2001 Cordylus langi – Frost, Janies, Mouton & Titus
2002 Cordylus langi – Anonymus/CITES

Loveridge, A. (1944): Revision of the African Lizards of the Family Cordylidae. — Bulletin of the Museum of Comparative Zoology (Cambridge, Mass.), 95 (1): 1–118. [type locality: “Mont-aux-Sources, Drakensberg, Basutoland”; type in MCZ]

Anonymus/CITES (2002): distr • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Bourquin & Channing 1980: distr, habitat  • Branch 1988: summary of data, conservation status “restricted”, map •  Branch 1998: morph, breeding, biology, habitat, distr, map  • Broadley 1964[b]: sys, morph, coloration, size, habitat, breeding, diet, distr  • Broadley 1971[b]: morph, distr  • Visser 1984[e]: morph, behav, distr •  Wermuth 1968: sys, distr  • van Wyk & Mouton 1992: epidermal glandular structure • 


Pseudocordylus melanotus melanotus (Smith, 1838) (Pseudocordylus melanotus (Smith, 1838 [Stanley et al. 2011])
1838 Cordylus (Pseudocordylus) melanotus Smith
1944 Pseudocordylus microlepidotus melanotus – Loveridge
1978 Pseudocordylus melanotus melanotus – de Waal
1984 Pseudocordylus melanotus melanotus – Visser
2001 Cordylus melanotus melanotus – Frost, Janies, Mouton & Titus
2002 Cordylus melanotus – Anonymus/CITES
2003 Cordylus melanotus melanotus – McConnachie & Whiting

Smith, A. (1838): Contributions to the South African Zoology. — Annals and Magazine of Natural History (London), (2) 2: 31–34. [as Cordylus (Pseudocordylus) melanotus; type locality: “Cape of Good Hope”, later given as Hills near Orange River east of “Phillopolis”, i.e. Phillipolis, Orange Free State (fide Loveridge 1944); type: lost]

Smith, 1838-1849
Click to enlarge


Anonymus/CITES (2002):
distr  • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Bates & Whittington-Jones 2009: distr Branch 1998: morph, breeding, biology, habitat, distr, map •  Cooper, van Wyk, Mouton, Al-Johany et al. 2000: antipredatory behavior  • Flemming 1992: reproductive cycle •  Flemming 1993[b]: reproductive cycle  • Flemming 1993[c]: reproductive cycle  •  Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces  • Loveridge 1944: revision (= all data) • McConnachie 2014: metabolic rate, ambush forager, energy budget  • McConnachie & Alexander 2004: digestive rate, body temperature effects, bomb calorimetry  • McConnachie, Alexander & Whiting 2007: thermal tolerance • McConnachie, Alexander & Whiting 2009 (2010): operative temperature, selected temperature, set-point range, sit-and-wait forager, thermoregulation •  McConnachie & Whiting 2003: tail loss, performance, antipredatory behaviour, tail energetic content •  Moon 2001: copulation, aggression  • Mouton & van Wyk 1993[a]: sexual dimorphism •  Mouton & van Wyk 1993[b]: distr, diet, reproduction, habitat  • Patterson & Bannister 1988: morph, behav  • Rogner 1994: morph, behav, ter •  Smith 1838-1849 [= 1843]: morph. = plate 25 (1843) fig. A (male) + B (female): Cordylus microlepidotus vars. = Pseudocordylus melanotus melanotus, fide Ulber 1999 •  Visser 1984[e]: morph, behav, distr •  de Waal 1978: sys, distr, for the OFS with map, morph, habitat, breeding, diet  • van der Wardt, Kik, Klaver, Janse & Beynen 1999: calcium balance •  Wermuth 1968: sys, distr  •


Pseudocordylus melanotus subviridis (Smith, 1838) (Pseudocordylus subviridis (Smith, 1838 [Stanley et al. 2011])
1838 Cordylus (Pseudocordylus) sub-viridis Smith
1937 Pseudocordylus microlepidotus subviridis – Fitzsimons
1943 Pseudocordylus subviridis subviridis – Fitzsimons
1984 Pseudocordylus melanotus subviridis – Visser
2001 Cordylus melanotus subviridis – Frost, Janies, Mouton & Titus
2002 Cordylus subviridis – Anonymus/CITES

Smith, A. (1838): Contributions to the South African Zoology. — Annals and Magazine of Natural History (London), (2) 2: 31–34. [as Cordylus (Pseudocordylus) sub-viridis; type locality: “Cape of Good Hope”, later given as Mountains behind Kaffirland and Natal (fide Loveridge 1944); type: lost]

Fitzsimons, 1943  Smith, 1838-1849
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Anonymus/CITES (2002): distr  • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Bourquin & Channing 1980: distr, habitat  • Branch 1981: distr, sys  • Branch & Burger 1991: predation by Lamprophis guttatus •  Broadley 1964[b]: sys, morph, coloration, size, habitat, breeding, diet, distr •  Broadley 1971[b]: morph, distr  • Fitzsimons 1937: type not found in NHM nor in Royal Scottish Museum  • Fitzsimons 1943: sys, morph, colour, size, distr, behav  • Fitzsimons 1948: sys, morph, distr •  Hewitt 1937[b]: morph, distr, behav •  Hughes 1988: age: >5 years  • Laforgia & Varano 1982: chromaffin cells in the adrenal glands  • Langerwerf 1996: behav, breeding in ter •  Langerwerf 2005: sys, distr, breeding in Alabama, USA  • Rose 1962: morph, behav  • Smet 1981: description of karyotypes •  Smith 1838-1849 [= 1843]: morph. = plate 26 (1843) fig. A (male), B (female) + C (young): Cordylus microlepidotus vars. = Pseudocordylus melanotus subviridis, fide Ulber 1999  • Switak 1980: behav, ter  • Visser 1984[e]: morph, behav, distr  • de Waal 1978: sys, distr, for the OFS with map, morph, habitat, breeding, diet  • Yeadon 1991[b]: size  •


Pseudocordylus microlepidotus microlepidotus (Cuvier, 1829)
1829 Cordylus microlepidotus Cuvier
1831 Zonurus microlepidotus – Gray
1834 Zonurus wittii Schlegel
1838 Cordylus (Pseudocordylus) montanus Smith
1943 Pseudocordylus microlepidotus microlepidotus – Fitzsimons
2001 Cordylus microlepidotus microlepidotus – Frost, Janies, Mouton & Titus
2002 Cordylus microlepidotus – Anonymus/CITES

Cuvier, G. (1829): Le Règne Animal distribué d’après son Organisation, pour servir de base à l’Histoire naturelle des Animaux et d’introduction à l’Anatomie Comparée. [2] Les Reptiles. — Paris: Déterville (2ième Édition), i–xvi + 1–406. [as Cordylus microlepidotus; type locality: “Africa” (fide Loveridge 1944); syntype in MHNP]

Fitzsimons, 1943  Smith, 1838-1849
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Anonymus/CITES (2002): distr  • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Böhme & Lang 1991: »Lacerta« rottensis von Meyer, 1856 to the Cordyliformes; compared with Pseudocordylus microlepidotus  • Boulenger 1885: morph •  Branch 1981: distr •  Branch 1998: morph, breeding, biology, habitat, distr, map •  Branch & Bauer 1995: behav  • Broadley 1971[b]: morph, distr  • Brygoo 1985[a]: sys, syntype in MHNP  • Cooper, van Wyk, Mouton, Al-Johany et al. 2000: antipredatory behavior  • Cooper, Whiting & van Wyk 1997: foraging modes •  Cunningham, Henderson & Tolley 2003: distr •  Fitzsimons 1943: sys, morph, colour, size, distr, behav •  Flower 1925[a]: age: >2 years •  Gibbons 1989: nematoda from Pseudocordylus micorlepidotus •  Gray 1845: morph •  Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces •  Hughes 1988: age: >2 years •  Loveridge 1944: revision (= all data)  • Mouton, Gagiano & Sachse 2005: generation glands, femoral glands •  Olmo, Capriglione, Odierna et al. 1995: genome characteristics  • Olmo & Odierna 1980: chromosomes, DNA, karyotype •  Rogner 1994: morph, behav, ter •  Rose 1962: morph, behav  • Schlegel 1834: morph, description of Zonurus wittii (type locality: “De zuidspits van Afrika”)  • Smith 1838: description of Cordylus (Pseudocordylus) montanus (type locality: “South Africa”) •  Smith 1838-1849 [= 1843]: morph. = plate 24 (1843) fig. 1: Cordylus microlepidotus = Pseudocordylus microlepidotus microlepidotus, fide Ulber 1999  • Snider & Bowler 1992: age: >2 years •  Visser 1984[e]: morph, behav, distr •  Wermuth 1968: sys, distr  • van Wyk & Mouton 1992: epidermal glandular structure • 


Pseudocordylus microlepidotus fasciatus (Smith, 1838)
1838 Cordylus (Pseudocordylus) fasciatus Smith
1838 Cordylus (Pseudocordylus) algoensis Smith
1927 Pseudocordylus fasciatus – Hewitt
1943 Pseudocordylus microlepidotus fasciatus – Fitzsimons
2001 Cordylus microlepidotus fasciatus – Frost, Janies, Mouton & Titus
2002 Cordylus fasciatus – Anonymus/CITES

Smith, A. (1838): Contributions to the South African Zoology. — Annals and Magazine of Natural History (London), (2) 2: 31–34. [as Cordylus (Pseudocordylus) fasciatus; type locality: “South Africa”; type: lost]

sc100  Smith, 1838-1849
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Anonymus/CITES (2002): distr  • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Branch 1981: distr •  Fitzsimons 1937: type not found in NHM nor in Royal Scottish Museum •  Fitzsimons 1943: sys, morph, colour, size, distr •  Hewitt 1937[b]: morph, distr, behav •  Loveridge 1944: revision (= all data) •  Smith 1838: description of Cordylus (Pseudocordylus) algoensis (type locality: “South Africa”) •  Smith 1838-1849 [= 1843]: morph. = plate 24 (1843) fig. 2: Cordylus microlepidotus = Pseudocordylus microlepidotus fasciatus, fide Ulber 1999 •  Smith 1838-1849 [1843]: morph.= plate 27 (1843) fig. 1: Cordylus fasciatus = Pseudocordylus microlepidotus fasciatus, fide Ulber 1999  • Wermuth 1968: sys, distr  •


Pseudocordylus microlepidotus namaquensis Hewitt, 1927
1927 Pseudocordylus microlepidotus namaquensis Hewitt
2001 Cordylus microlepidotus namaquensis – Frost, Janies, Mouton & Titus

Hewitt, J. (1927[a]): Further descriptions of reptiles and batrachians from South Africa. — Rec. Albany Museum (Grahamstown), 3 (5): 371–415. [type locality: “Namaqualand”; type in South African Museum, Cape Town]

sc_162

Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Branch 1981: distr •  Fitzsimons 1943: sys, morph, colour, size, distr •  Loveridge 1944: revision (= all data)  • Wermuth 1968: sys, distr  •


Pseudocordylus nebulosus Mouton & van Wyk, 1995 (Hemicordylus nebulosus (Mouton & van Wyk, 1995 [Stanley et al. 2011])
1995 Pseudocordylus nebulosus Mouton & van Wyk
2001 Cordylus nebulosus – Frost, Janies, Mouton & Titus
2002 Cordylus nebulosus – Anonymus/CITES

Mouton, P. & van Wyk, J. (1995): A new crag lizard from the Cape Folded Mountains in South Africa. — Amphibia-Reptilia (Leiden), 16 (4): 389–399. [type locality: “Northern slope of Landdroskop in the Hottentotsholland Mountains, in the Western Cape, South Africa, at an altitude of 1200m above sea level”; type in South African Museum, Cape Town]

Anonymus/CITES (2002): distr  •  Baard & de Villiers 2000: conservation • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Branch 1998: morph, breeding, biology, habitat, distr, map • Costandius & Mouton 2006: epidermal glands, body size, head size, melanistic, fecundity selection 


Pseudocordylus spinosus Fitzsimons, 1947
1947 Pseudocordylus spinosus Fitzsimons
2001 Cordylus spinosus – Frost, Janies, Mouton & Titus
2002 Cordylus spinosus – Anonymus/CITES

Fitzsimons, V. (1947): Descriptions of New Species and Subspecies of Reptiles and Amphibians from Natal, together with Notes on some other Little Known Species. — Annals of the Natal Museum (Pietermaritzburg), 11 (1): 111–137. [see below]  [type locality: “Cathkin Peak Area, Drakensberg, Natal”; type in  TM]

Fitzsimons, 1947


Anonymus/CITES (2002): distr  • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Bourquin & Channing 1980: distr, habitat  • Branch 1988: summary of data, conservation status “restricted”, map •  Branch 1998: morph, breeding, biology, habitat, distr, map  • Broadley 1964[b]: sys, morph, coloration, size, habitat, breeding, diet, distr  • Broadley 1971[b]: morph, distr  • Hughes 1988: age: >1 year  • Visser 1984[e]: morph, behav, distr •  de Waal 1978: sys, distr, for the OFS with map, morph •  Wermuth 1968: sys, distr  •


Pseudocordylus transvaalensis Fitzsimons, 1943
1943 Pseudocordylus subviridis transvaalensis Fitzsimons
1984 Pseudocordylus melanotus transvaalensis – Visser
1998 Pseudocordylus transvaalensis – Branch
2001 Cordylus transvaalensis – Frost, Janies, Mouton & Titus
2002 Cordylus transvaalensis – Anonymus/CITES

Fitzsimons, V. (1943): The Lizards of South Africa. — Memoirs of the Transvaal Museum (Pretoria), No. 1, xv + 528 pp. [as Pseudocordylus subviridis transvaalensis; type locality: “Woodbush, Pietersburg District, North Transvaal”; type in TM]

Anonymus/CITES (2002): distr  • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Branch 1998: morph, breeding, biology, habitat, distr, map •  Broadley 1964[b]: sys, morph, coloration, size, habitat, breeding, diet, distr  • Visser 1984[e]: morph, behav, distr •  de Waal 1978: sys • Wever 1978: structure and function of ear •  Yeadon 1991[a]: size, distr • 


© Klaus Adolphs  – created: 26 February 2001  – last changed or updated: 20 July 2014