This Gerrhosaurus+ site includes also the genera Broadleysaurus and Matobosaurus.

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Wiegmann, A. (1828):
Beyträge zur Amphibienkunde. — Isis von Oken (Leipzig), 21: 364–383. [type species: Gerrhosaurus flavigularis Wiegmann, 1828 by monotypy] •  molecular phylogeny, all species (!): Bates et al. 2013.

Distribution: Africa, south of 5 deg. northern latitude, except Zaire (forest areas) and Kalahari. [Lang 1991: map]


Gerrhosaurus auritus Boettger, 1887
1887 Gerrhosaurus auritus Boettger
1942 Gerrhosaurus nigrolineatus auritus – Loveridge
1988 Gerrhosaurus multilineatus auritus – Branch
2003 Gerrhosaurus auritus – Griffin
2007 Gerrhosaurus auritus – Broadley (in book review)
2013 Gerrhosaurus auritus – Bates et al.

Boettger, O. (1887): Zweiter Beitrag zur Herpetologie Südwest- und Süd-Afrikas. — Bericht über die Senckenbergische naturforschende Gesellschaft in Frankfurt am Main (Frankfurt), (1886/1887), 135–173. [see below] [as Gerrhosaurus auritus; type locality: “Ondonga, Ovamboland, Namibia”; type in SMF]

Boettger, 1887Fitzsimons, 1943
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Auerbach 1985: morph, behav  • Auerbach 1986: etymology •  Auerbach 1987: morph, behav, distr • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Bowler 1977: age: >8 years •  Fitzsimons 1935[b]: distr, size, morph, colour  • Fitzsimons 1943: sys, morph, colour, size, distr, behav •  Griffin 2003: conservation status, distr  • Hewitt 1910[b]: morph, distr •  Hughes 1988: age: >8 years •  Loveridge 1942[a]: revision (=all data)  • Mertens 1922: type in SMF •  Mertens 1955[b]: sys, distr  • Mertens 1967: type in SMF •  Rose 1962: morph, behav •  Snider & Bowler 1992: age: >8 years •  Visser 1984[a]: morph, behav, distr •  Wermuth 1968: sys, distr  •


Gerrhosaurus bulsi Laurent, 1954
1954 Gerrhosaurus auritus bulsi Laurent
2004 Gerrhosaurus bulsi – Broadley & Cotterill
2007 Gerrhosaurus bulsi – Broadley (in book review)
2013 Gerrhosaurus bulsi – Bates et al.

Laurent, R. F. (1954): Reptiles et Batraciens de la region de Dundo (Angola) (Deuxieme note). – Com. Dia. Angola, Ser. Cul. Publ. Cult., 23: 37-84. [type locality: “Dundo, Angola”]

Gerrhosaurus bulsi

Wagner 2010: taxonomy, description, distribution, habitat


Gerrhosaurus flavigularis Wiegmann, 1828
1834 Gerrhosaurus ocellatus Cocteau
1836 Pleurotuchus desjardinii Smith
1836 Pleurotuchus chrysobronchus Smith
1844 Gerrhosaurus bibroni Smith
1883 Gerrhosaurus flavigularis var. quadrilineata Boettger
1942 Gerrhosaurus flavigularis flavigularis – Loveridge
1942 Gerrhosaurus flavigularis fitzsimonsi Loveridge
1991 Gerrhosaurus flavigularis – Broadley & Howell
1998 Gerrhosaurus flavigularis – Branch
2013 Gerrhosaurus flavigularis – Bates et al.

Wiegmann, A. (1828): Beyträge zur Amphibienkunde. — Isis von Oken (Leipzig), 21: 364–383. [type locality: “Africa merid.”; syntype in ZMB]

Gerrhosaurus flavigularis (c) Ingo Kober  Fitzsimons, 1943  Smith, 1838-1849  Smith, 1838-1849
Picture-Taker-hQbzH5  Picture-Taker-XkI7c7
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Alexander 1990:
distr •  Auerbach 1985: morph, behav  • Auerbach 1986: etymology •  Auerbach 1987: morph, behav, distr • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Barrio & Santoma 1996: ter •  Bauer 2000: about type locality, sys  • Bauer, Lang & Günther 1994: sys, about type •  Beddaed 1905[a]: arterial system •  Beddard 1905[b]: anatomy •  Berger-Dell’mour 1987: distr •  Boettger 1883: description of Gerrhosaurus flavigularis var. quadrilineata (type locality: “Smithfield, Transvaal”, type in SMF) •  Böhme 1988: genital morphology  • Boulenger 1887[a]: morph •  Bourquin & Channing 1980: distr, habitat  • Boycott & Morgan 1988: reproduction  • Branch 1976: Gerrhosaurus as stomach contents of Lycophidion capense  • Branch 1998: morph, breeding, biology, habitat, distr, map •  Branch 2005: morph, distr, photo  • Broadley 1959[b]: morph, behav  • Broadley 1962[a]: morph, size, habitat  • Broadley 1971[c]: distr, habitat  • Broadley & Howell 1991: sys, distr  • Brygoo 1985[a]: sys, syntypes of Gerrhosaurus bibroni and Gerrhosaurus ocellatus in MHNP •  Clauss & Clauss 2002: distr, biology •  Cocteau 1834: description of Gerrhosaurus ocellatus, type locality: “Cap de Bonne Espérance” •  Cope 1862: morph •  Cott 1934: morph, colour, behav  • Douglas 1992: reproduction  •  Fitzsimons 1935[b]: distr, morph, size, colour  • Fitzsimons 1937: about type of Gerrhosaurus bibroni •  Fitzsimons 1939[b]: morph, colour, size •  Fitzsimons 1943: sys, morph, colour, size, distr, behav •  Flower 1937: age: >11 years  • Gray 1845: morph •  Griffin 2003: conservation status, distr  • Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces  • Hedges 1983: behav •  Hewitt 1910[b]: morph, distr  • Hewitt 1937[b]: morph, distr, behav •  Hughes 1988: age: >11 years •  Girard & Plazanet 2002: autotomy  • Kober 1998 and 1999: behav •  Kober 1990: ter, breeding  • Largen & Spawls 2010: description, photo, distribution, map, ecology •  Loveridge 1920: morph, behav  • Loveridge 1942[a]: revision (=all data) and description of Gerrhosaurus flavigularis fitzsimonsi (type locality: “Mt. Mbololo, Teita, Kenya Colony”; type in MCZ) •  Loveridge 1953[a]: morph, size, breeding •  Malan 1941: cranial anatomy •  Manacas 1952: morph •  Mitchell & Steyn 1965: morph, colour, distr  • Nieden 1913: morph •  Nietzke 1998: ter, morph, distr, behav  • Nietzke 1978: ter, morph  • Odierna et al. (2002): Karyotypes •  Patterson & Bannister 1988: morph, behav  • Peters 1882: morph, distr  • Pienaar 1978: morph, distr, behav •  Pienaar, Haacke & Jacobsen 1983: morph, distr, behav •  Popp 1958: ter  • Rogner 1994: morph, behav, ter •  Rose 1962: morph, behav  • Schmidt 1998: ter •  Smet 1981: description of karyotypes •  Smith 1836: description of Pleurotuchus Desjardinii (type locality: “country alongst the south-east coast”) and of Pleurotuchus chrysobronchus (type locality: “towards the sources of the Cowie river, about Graham’s Town and Blue Kranz”) •  Smith 1838-1849 [= 1844]: description of Gerrhosaurus bibroni (type locality: “towards the sources of the Caledon, one of the principal branches of the Orange River”, type in NHM) = plate 38 (1844) fig. 1: Gerrhosaurus bibroni •  Smith 1838-1849 [= 1844]: morph • Snider & Bowler 1992: age: >1 year  • Spawls, Howell, Drewes & Ashe 2001: morph, distr, map, habitat, biology •  Stebbins 1961: body temperature  • Tapley, Griffiths & Bride 2011: trade in UK •  Tornier 1902: distr, morph  • de Waal 1978: sys, distr, for the OFS with map, morph, habitat, breeding, diet • Wever 1978: structure and function of ear •  Wermuth 1968: sys, distr  • Werner 1910: morph, distr  •



Gerrhosaurus intermedius Lönnberg, 1907
1907 Gerrhosaurus flavigularis forma intermedia Lönnberg
2013 Gerrhosaurus intermedius – Bates et al.

For more details see: Bates et al. 2013



Gerrhosaurus major major Duméril, 1851 (Broadleysaurus major (Duméril, 1851) [Bates et al., 2013])
1889 Gerrhosaurus zanzibaricus Pfeffer
1906 Gerrhosaurus bergi Werner
1908 Gerrhosaurus grandis Boulenger
1910 Gerrhosaurus major var. grandis – Hewitt
1987 Gerrhosaurus major major – Broadley
2013 Broadleysaurus major – Bates et al.

Duméril, A. (1851): Catalogue Méthodique de la Collection des Reptiles. — Paris: Gide et Baudry, iv + 224 pp. [type locality: “Zanzibar”; syntype in MHNP]

Gerrhosaurus major  Boulenger, 1908
 Fitzsimons, 1943  Werner, 1906
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Auerbach 1987:
morph, behav, distr •  Barrio & Santoma 1996: ter • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Bech & Kaden 1990: ter •  Bellekom 1990: ter •  Böhme 1975: distr, sexualdichromatism •  Böhme 1988: genital morphology •  Boulenger 1887[a]: morph •  Boulenger 1908[a]: description of Gerrhosaurus grandis (type locality: “Ubombo, Zululand”, type in NHM)  • Bowker 1984: thermoregulatory abilities, patterns of thermoregulation •  Bowler 1977: age: >11 years  • Boycott & Morgan 1988: reproduction  •  Branch 1998: morph, breeding, biology, habitat, distr, map •  Branch 2005: morph, distr, photo  • Broadley 1959[b]: morph, behav  • Broadley 1962[a]: morph, size, habitat, diet  • Broadley 1971[b]: morph, distr •  Broadley 1971[c]: distr, habitat  • Broadley 1987: review of variations, sys, map  • Brygoo 1985[a]: sys, about syntype  • Cobron 1987: morph, distr, habitat, ter  • Cooper, van Wyk, Mouton, Al-Johany et al. 2000: antipredatory behavior  • Cott 1934: colour, behav •  Couvering 1979: fossil Gerrhosaurus • Dassow 2008: ter, breeding, diet  • Dathe 1987: morph, behav, ter •  Dijk, Manley & Gallo 1998: spontaneous otoacoustic emissions •  Don, Estany, Ramos et al. 1994: fever  • Estes 1962: fossil Gerrhosaurus •  Evans 1994: morph, ter, diet, breeding  • Fitzsimons 1939[b]: morph, colour, size •  Fitzsimons 1943: sys, morph, colour, size, distr, behav •  Flower 1937: age: >7 years •  Haagner & Morgan 1992: behav, courtship, mating, reproduction, hatching  • Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces  • Hedges 1983: behav  • Hewitt 1910[b]: morph, distr  • Hughes 1988: age: >16 years  • John 1980: behav, morph, habitat, ter •  Kellough 1989: ter, breeding •  Laforgia & Varano 1982: chromaffin cells in the adrenal glands • Largen & Spawls 2010: description, photo, distribution, map, ecology   Loveridge 1920: morph, behav  • Loveridge 1942[a]: revision (= all data) •  Loveridge 1953[a]: morph, size, breeding, habitat  • Montuelle et al. 2009: feeding • Muchlinski, Estany, Salguero et al. 1998: acute phase response •  Muchlinski, Gramajo & Garcia 1999: antibiotic, body temperature set-point, fever, telemetry, temperature regulation  • Nieden 1913: morph •  Nietzke 1998: ter, morph, behav, diet, reproduction •  Nietzke 1978: ter, morph  • Odierna et al. (2002): Karyotypes  • Olmo, Capriglione, Odierna et al. 1995: genome characteristics  • Olmo & Odierna 1980: chromosomes, DNA, karyotype •  Patterson & Bannister 1988: morph, behav  • Peters 1882: morph, distr  • Pfeffer 1889: description of Gerrhosaurus zanzibaricus (type locality: “Zanzibar, Kibueni”)  • Pienaar 1978: morph, distr, behav •  Pienaar, Haacke & Jacobsen 1983: morph, distr, behav •  Rese 1986: morph, distr, ter, reproduction, diet, behav •  Rogner 1994: morph, behav, ter •  Rowe-Rowe 1968: ter, diet, behav  •  Schmidt 1998: ter •  Smet 1981: description of karyotypes •  Snider & Bowler 1992: age: >24 years  • Spawls, Howell, Drewes & Ashe 2001: morph, distr, map, habitat, biology • Tapley, Griffiths & Bride 2011: trade in UK • Tornier 1902: distr, morph • Visser 1984[a]: morph, behav, distr • Wever 1978: structure and function of ear •  Wermuth 1968: sys, distr  • Werner 1906: description of Gerrhosaurus bergi (type locality: “Usambara”, holotype in ZMB)  •


Gerrhosaurus major bottegoi del Prato, 1895 = (Broadleysaurus major (Duméril, 1851) [Bates et al., 2013])
1895 Gerrhosaurus bottegoi del Prato
1901 Gerrhosaurus major var. zechi Tornier
1919 Gerrhosaurus zechi – Schmidt
1987 Gerrhosaurus major bottegoi – Broadley
2013 Broadleysaurus major – Bates et al.

Prato, A. del (1895): Vertebrati Eritri aggiunta al catalogo della collezione Eritrea Bottego. — Atti. Soc. Ital. Sci. Nat. (Milano), 35: 17–26. [see below] [as Gerrhosaurus bottegoi; type locality: “Valle di Ghinda, Eritrea”; type in Museo Bettego, Parma]

del Prato, 1895  Schmidt, 1919
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Bates, Branch, Bauer et al. (Atlas and Red List) 2014:
tax, habitat, map, photo, conservation • Bauer, Lang & Günther 1994: sys, about syntype of Gerrhosaurus major zechi in ZMB  • Broadley 1987: review of variations, sys, map  • Chirio & LeBreton 2007: as Gerrhosaurus major zechi, incl. foto •  Daszak & Ball 2001: description of parasites  • Dunger 1967: morph, behav, distr, size •  Hedges 1983: behav • Ineich 1999/2001: Gerrhosaurus major bottegoi in Djibouti •  Loveridge 1933: morph, diet, habitat  • Loveridge 1942[a]: revision (=all data)  • Parker 1942: morph, colour, sys  • Schmidt 1919: morph, distr, colour, behav  •   Tornier 1901: description of Gerrhosaurus major var. zechi (type locality: “Kete Kratje, Togo”)  • Trape, Trape & Chirio (2012): description, distribution map, natural history, fotos •  Wermuth 1968: sys, distr  •


Gerrhosaurus multilineatus Bocage, 1866
1866 Gerrhosaurus multilineatus Bocage
2007 Gerrhosaurus multilineatus – Broadley (in book review)
2013 Gerrhosaurus multilineatus – Bates et al.

Bocage, J. (1866[a]): Lista dos reptis das possessões portuguezas d’Africa occidental que existem no Museu de Lisboa. — Jornal de Sciencias Mathematicas, Physicas e Naturaes (Lisboa), 1: 37–56. [type locality: “Duque de Braganca, Angola”; type in Museu de Lisboa]  — and  [this paper follows the before-mentioned:] Bocage, J. (1866[b]): Reptiles nouveaux ou peu connus recueilles dans les possessions portugaises de l’Afrique occidentale, qui se trouvent au Muséum de Lisbonne. — Jornal de Sciencias Mathematicas, Physicas e Naturaes (Lisboa), 1: 57–78.

Branch 1998: morph, breeding, biology, habitat, distr, map •  Broadley 1960: description of Gerrhosaurus nigrolineatus anselli (type locality: “Mwinilunga Boma, Northern Rhodesia”, type in National Museum Southern Rhodesia) •  Broadley 1971[c]: distr, habitat  • Broadley 1991: habitat, reproduction •  Broadley & Cotterill 2004: distr, sys • Haagner, Branch & Haagner 2000: size, diet, reproduction, parasites, as Gerrhosaurus bulsi, with a note that “the taxon G. multilineatus Bocage is based on a hybrid specimen. The name is therefore unavailable.” • Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces •  Laurent 1954[a]: description of Gerrhosaurus auritus bulsi, type locality: “Dundo”)  • Laurent 1964: morph, sys  •  


Gerrhosaurus nigrolineatus nigrolineatus Hallowell, 1857
(1907 Gerrhosaurus flavigularis forma intermedia Lönnberg – full species, see above)
1919 Gerrhosaurus flavigularis nigrolineatus – Schmidt
1939 Gerrhosaurus nigrolineatus australis Fitzsimons
1968 Gerrhosaurus nigrolineatus nigrolineatus – Wermuth
1998 Gerrhosaurus nigrolineatus – Branch
2013 Gerrhosaurus nigrolineatus – Bates et al.

Hallowell, E. (1857): Notice of a collection of Reptiles from the Gaboon country, West Africa, recently presented to the Academy of Natural Sciences of Philadelphia, by Dr. Henry A. Ford. — Proceedings of the Academy of Natural Sciences of Philadelphia, 9 (1857): 48–72. [type locality: “Gaboon”; type in Academy of Natural Sciences of Philadelphia]

Gerrhosaurus nigrolineatus  Fitzsimons, 1943  Hellmich & Schmelcher, 1956
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Auerbach 1986: etymology •  Auerbach 1987: morph, behav, distr •  Boettger 1888: morph, distr •  Boulenger 1887[a]: morph •  Bowker 1984: thermoregulatory abilities, patterns of thermoregulation •  Branch 1976: Gerrhosaurus as stomach contents of Lycophidion capense  • Branch 1998: morph, breeding, biology, habitat, distr, map •  Branch 2005: morph, distr, photo  • Broadley 1959[b]: morph, behav  • Broadley 1962[a]: morph, size, habitat, diet  • Broadley 1971[c]: distr, habitat •  Broadley & Cotterill 2004: distr •  Cooper 1992: tongue flicking •  Cooper & Habegger 2001: behav, omnivory, food chemical discrimination •  Cooper & Trauth 1992: pheromone, cloacal glands, tongue-flicks  • Cope 1896: hemipenis •  Fitzsimons 1935[b]: distr, morph, size, colour  •  Fitzsimons 1938: morph, colour •  Fitzsimons 1939[a]: description of Gerrhosaurus nigrolineatus australis (type locality: “Kaapmuiden, Eastern Transvaal”, type in TM)  • Fitzsimons 1939[b]: morph, colour, size •  Fitzsimons 1943: sys, morph, colour, size, distr, behav • Goldberg & Bursey 2010: endoparasites • Goldberg et al. 2014: endoparasites • Griffin 2003: conservation status, distr  • Haagner & Morgan 1991: reproduction  • Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces •  Hellmich 1954-55: behav •  Hellmich 1957[a]: distr, morph  •  Hellmich 1957[b]: distr, morph, behav •  Hellmich & Schmelcher 1956: description of Gerrhosaurus nigrolineatus ahlefeldti (type locality: “Piri-Dembos, Nova Roca Douro”; type in Zoological Museum, Hamburg)  • Huff 1989: ter •  Lönnberg 1907: description of Gerrhosaurus flavigularis forma intermedia (type locality: ”Steppe near Natron Lake, Kibonoto, Tanganyika”)  • Loveridge 1920: morph, behav  • Loveridge 1923[b]: morph •  Loveridge 1942[a]: revision (= all data)  • Loveridge 1942[b]: breeding  • Loveridge 1953[a]: morph, size •  Manacas 1963: morph, colour •  Mertens 1955[b]: morph, distr  • Mertens 1970: age: >4 years •  Nietzke 1978: ter, morph  •  Peterson & Bezy 1985: microstructure of scale surfaces  • Pienaar 1978: morph, colour, size, distr, behav  • Pienaar, Haacke & Jacobsen 1983: morph, colour, distr, behav •  Popp 1958: ter •  Rogner 1994: morph, behav, ter •  Schmidt 1919: morph, distr, behav •  Schmidt 1913: skin, dermal bones  • Spawls, Howell, Drewes & Ashe 2001: morph, distr, map, habitat, biology •  Sternfeld 1912: morph  • Versluys 1898: description of the middle and outer ear •  Visser 1984[a]: morph, behav, distr •  Wermuth 1968: sys, distr  • Wever 1970: fine structure of the ear • Wever 1978: structure and function of ear •


(?) Gerrhosaurus nigrolineatus ahlefeldti  Hellmich & Schmelcher, 1956
status not clear

Franzen & Glaw 2007: remarks on the types  •


Gerrhosaurus skoogi Andersson, 1916
1916 Gerrhosaurus skoogi Andersson
1942 Gerrhosaurus validus skoogi – Loveridge
1953 Angolosaurus skoogi – Fitzsimons
2003 Gerrhosaurus skoogi – Griffin
2003 Gerrhosaurus skoogi – Lamb, Meeker, Bauer & Branch
2007 Angolosaurus skoogi – Nance
2013 Gerrhosaurus skoogi – Bates et al.

Andersson, L. (1916): Notes on the reptiles and batrachians in the Zoological Museum at Gothenburg with an account of some new species. — Göteborgs Kungliga Vetenskap och Vitter Hets-Samnalles Handlingar Sjatte Foljden, Series B, 4, (17, No. 5) [= Meddelanden fran Göteborgs Musei Zool. Afdel. (Goteborg), No. 9]: 1–41. [as Gerrhosaurus skoogi; type locality: “Port Alexander, Portuguese West Africa”; type in Göteborg Museum] Distribution: coastal zone of north-west Namibia, south-west Angola (Namib desert). [Lang 1991: map]

Gerrhosaurus skoogi


Arnold 1995: history of sand diving in the Gerrhosauridae (Angolosaurus) •  Branch 1998: morph, breeding, biology, habitat, distr, map •  Broadley 1971[b]: morph, distr  • Chinsamy, Hanrahan, Neto & Seely (1995): skeletochronological assessment of age in Angolosaurus •  Clarke & Nicolson 1994: water, energy and electrolyte balance •  Cooper 1995: “...also readily consumes insects in the laboratory” •  Fitzsimons 1953: new genus (Angolosaurus), morph, size, colour, behav, sys  • Griffin 2003: conservation status, distr  • Hamilton & Coetzee 1969: thermoregulatory behaviour  • Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces  • Lamb, Meeker, Bauer & Branch 2003: DNA sequence analysis, systematic status, replace of Angolosaurus within Gerrhosaurus •  Loveridge 1942[a]: revision (= all data) •  Mitchell, Seely, Roberts et al. 1987: morph, population characteristics, feeding, predation, behav, locomotion, thermoregulation, metabolic rate •  Nagy, Clarke, Seely et al. 1991: water and energy balance  • Nance 2007: anatomy, skull, mandible  •  Pietruszka 1988[b]: sexual dimorphism, social structure •  Pietruszka, Hanrahan, Mitchell et al. 1986: diet •  Rogner 1994: morph, behav, ter •  Seely, Mitchell & Goelst 1990: thermoregulatory behaviour  • Seely, Mitchell & Roberts et al. 1988: behav •  Steyn 1963: distr, morph, behav, diet •  Visser 1984[b]: morph, behav, distr •  Wermuth 1968: sys, distr  •


Gerrhosaurus typicus (Smith, 1836)
1836 Pleurotuchus typicus Smith
1839 Gerrhosaurus typicus – Duméril & Bibron
2013 Gerrhosaurus typicus – Bates et al.

Smith, A. (1836): The Characters of two new Genera of South African Reptiles with Descriptions of Species belonging to each. — Magazine of Zoology and Botany (Jardine), (Edinburg, London & Dublin), [1837], 1 (2): 141–145. [as Pleurotuchus typicus; type locality: “dry sandy flats of Little Namaqualand, and is rather abundant towards the mouth of the Orange river”; type in NHM]

Fitzsimons, 1943  Smith, 1838-1849
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Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Boulenger 1887[a]: morph •  Branch 1998: morph, breeding, biology, habitat, distr, map •  McLachlan 1988: summary of data, conservation status: “rare”, map •  Branch & Bauer 1995: behav •  Fitzsimons 1937: type in NHM •  Fitzsimons 1943: sys, morph, colour, size, distr, behav •  Gray 1845: morph •  Griffin 2003: conservation status, distr  • Hughes 1988: age: >2 years •  Loveridge 1942[a]: revision (=all data)  • Malan 1941: cranial anatomy •  Smith 1838-1849 [= 1844]: morph. plate 38 fig. 2 •  Visser 1984[a]: morph, behav, distr •  Wermuth 1968: sys, distr  •


Gerrhosaurus validus validus Smith, 1849 (Matobosaurus validus (Smith, 1849) [Bates et al., 2013])
1854 Gerrhosaurus robustus Peters
1930 Gerrhosaurus ciprianii Scortecci
1942 Gerrhosaurus validus validus – Loveridge
2013 Matobosaurus validus – Bates et al.

Smith, A. (1838–1849) [= 1849]: Illustrations of the Zoology of South Africa. Consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836. Fitted out by ‘The Cape of Good Hope Association for Exploring Central Africa’. — London: Smith, Elder and Co., [Appendix] [as Gerrhosaurus vallidus; type locality: “Sources of the Garrep or Orange River, South Africa”; type in NHM]

Gerrhosaurus validus  Fitzsimons, 1943  Peters, 1882Picture-Taker-G2hDgR
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Auerbach 1986: etymology •  Auerbach 1987: morph, behav, distr •  Barrio & Santoma 1996: ter • Bates, Branch, Bauer et al. (Atlas and Red List) 2014: tax, habitat, map, photo, conservation • Bauer, Lang & Günther 1994: sys, about type •  Biegler 1966: age: >10 years  •  Bissinger & Simon 1979: tongue extrusions •  Boulenger 1887[a]: morph •  Bowler 1977: age: >4 years  • Boycott & Morgan 1988: reproduction  • Branch 1998: morph, breeding, biology, habitat, distr, map •  Branch, Rödel & Marais 2005: distr Broadley 1959[b]: morph, behav  • Broadley 1962[a]: morph, size, habitat, diet  • Broadley 1971[b]: morph, distr  • Broadley 1971[c]: distr, habitat  •  Broom 1935: skull •  Cooper, van Wyk, Mouton, Al-Johany et al. 2000: antipredatory behavior  • Cooper, Whiting & van Wyk 1997: foraging modes  • Cooper & Steele 1999: foraging and chemosensory behavior  • Cooper 2000: chemosensory response to food •  Falk 1921: behav, ter  • Fitzsimons 1935[b]: distr, size, morph, colour  • Fitzsimons 1937: type in NHM •  Fitzsimons 1939[b]: morph, colour, size •  Fitzsimons 1943: sys, morph, colour, size, distr, behav •  Griffin 2003: conservation status, distr  • Harbig (2003): captive care, breeding raising of juveniles, social interactions •  Harvey & Gutberlet 1995: microstructure, evolution and ontogeny of scale surfaces  • Hering-Hagenbeck, Petter & Boomker 2002: description of parasites  • Hewitt 1910[b]: morph, distr  •  Hughes 1988: age: >10 years  • Kellough 1989: ter, breeding •  Köhler 1990: ter, breeding •  Laforgia & Varano 1982: chromaffin cells in the adrenal glands  • Loveridge 1942[a]: revision (= all data)  • Loveridge 1953[a]: morph, size, breeding, habitat  • Manacas 1952: morph, colour •  Olmo, Capriglione, Odierna et al. 1995: genome characteristics  • Olmo & Odierna 1980: chromosomes, DNA, karyotype •  Patterson & Bannister 1988: morph, behav  • Peters 1854: description of Gerrhosaurus robustus, type locality: “Tette”, lectotype in ZMB •  Peters 1882: morph, sys, distr  • Pienaar 1978: colour, size distr, behav •  Pienaar, Haacke & Jacobsen 1983: colour, distr, behav  • Rogner 1994: morph, behav, ter •  Schmidt 2005: Haltung, Zucht im Familienverband •  Schmidt 2004: ter, breeding •  Schmidt 2008: ter, breeding •  Scortecci 1930[b]: description of Gerrhosaurus ciprianii (type locality: “Old Enchopi, Dintorni di Mozambico”, type in [?]Museo Civico di Storia Nat. di Milano)  • Snider & Bowler 1992: age: >16 years  • Steinmetz, Pütsch & Bisschopinck 1998: transport mortality  • Switak 1979: behav, ter  • Switak 1988: behav, ter  • Ulber 1999: about the spelling „vallidus“  • Visser 1984[a]: morph, behav, distr •  Wermuth 1968: sys, distr  • Wever 1978: structure and function of ear •


Gerrhosaurus validus maltzahni de Grys, 1938 (Matobosaurus maltzahni (de Grys, 1938) [Bates et al., 2013])
1938 Gerrhosaurus maltzahni de Grys
1938 Gerrhosaurus validus damarensis Fitzsimons
1942 Gerrhosaurus validus maltzahni – Loveridge
2013 Matobosaurus maltzahni – Bates et al.

Grys, P. de (1938): Gerrhosaurus Maltzahni spec. nov. — Zoologischer Anzeiger (Leipzig), 124 (3/4): 58–60. [see below] [as Gerrhosaurus maltzahni; type locality: “Farm Roidina, D.S.W.-Afrika”; type in Zoological Museum Hamburg]

de Grys, 1938  Fitzsimons, 1938 
photo by Johan Marais  photo by Johan Marais 
Click to enlarge


Fitzsimons 1938: description of Gerrhosaurus validus damarenis, type locality: “Kamanyab”, type in TM  • Fitzsimons 1943: sys, morph, colour, size, distr, behav  • Hellmich 1954-55: behav •  Hellmich 1957[a]: distr, morph  • Hellmich 1957[b]: distr, morph, behav •  Hoffmann 1989: distr •  Loveridge 1942[a]: revision (=all data)  • Mertens 1955[b]: sys •  Rose 1962: morph, behav  • Visser 1984[a]: morph, behav, distr •  Wermuth 1968: sys, distr  •



© Klaus Adolphs  – created: 26 February 2001  – last changed or updated: 29 July 2015